Growth Form:
Evergreen shrub or treelet usually to 4.5(–10) m tall. plants vary from low shrubs to trees with open, wide–branched crowns.
Stems:
Bark often whitish, young stems usually pubescent.
Roots:
Leaves:
Leaves simple.
Alternate.
Blades obovate–oblong to oblanceolate, 5–12 cm long x 2–3.5 cm wide.
Apex rounded or slightly notched.
Upper surfaces glabrous, dark green; lower surfaces pubescent ± whitish gray; blades papery to leathery.
Margins entire or apex weakly toothed, rarely with a few small teeth in the upper half.
Petiolate.
Stipules minute.
Flowers:
Flowers in stalked clusters in leaf axils or along stem between leaves; staminate flowers to 15 per cluster; pistillate flowers 1–5 per cluster; peduncles flattened, 1.3–5 cm long.
Flowers unisexual, ca 5 mm in diameter. pedicels ca. 2–6 mm long with 2 bracteoles below the middle, these 2–3 mm long.
Sepals persistent.
Petals white or greenish, usually as many as sepals; rotate, connate at base, caducous.
Stamens as many as petals, inserted at the base of the corolla, staminodia in pistillate flowers similar to fertile stamens, but usually smaller.
Ovary superior; 4-celled.
Fruit:
Fruit red to yellow in stalked clusters; globose; 6–7 mm in diameter; stones 1–4; ribbed.
Seeds with abundant oily or proteinaceous endosperm.
Ploidy:
Habitat:
Elevation Range:
Historical Distribution
Uses and Culture
USES
Natural History
Statewide Status
Naturalized
Island Status
O'ahu Naturalized
Hawai'i Naturalized
Dispersal Agents
Water/Ocean
Birds
Pollinators
Specimens
Conservation Status
IUCN:Least Concern
Hawaii Pacific Weed Risk
Assessment Rating:High Risk
HPWRA Score:9
Notes
Description digitized by Pumehana Imada
Native to the Atlantic Coastal Plain of the southeastern and eastern US and reported from Cuba and the Bahama Islands. Planted along highway from Hilo to Volcano, Hawai‘i and in Honolulu, O‘ahu and now reported to be naturalizing [Imada, et al].
Bibliography
Name Published In: Sp. Pl.: 125 (1753)
Other References
Wagner et al. 1990:222 (O?, H? [planted but not nat.]); Imada et al. 2000:9 (NEWNAT/O, H); Staples & Herbst 2005:129 (KEY), 130 (DESCR)
Wagner, W. L., Herbst, D. R., & Sohmer, S. H. (1999). Manual of the Flowering Plants of Hawai'i, Vols. 1 and 2 (No. Edn 2). University of Hawai'i and Bishop Museum Press.
Staples, G., & Herbst, D. R. (2005). A Tropical Garden Flora. Bishop Museum Press.
Occurrences
SNo.
Scientific Name
Locality
Habitat
Basis of Record
Recorded By
Record Number
Island
Source
Date
1
Ilex cassine
Schofield Barracks, East Range
Eroded slope on edge of flat, grassy, alien-dominated ridge.
Preserved_Specimen
Kawelo, K.
14
Oahu
BISH
2005-04-21
2
Ilex cassine
Kakaako
Street tree
Preserved_Specimen
Allison, A.
Oahu
BISH
2021-06-01
3
Ilex cassine
Helemano Forest Reserve above NCTAMS
Mixed alien and native forest above NCTAMS military reservation
Preserved_Specimen
Ching-Harbin, S.
2020120301
Oahu
BISH
2020-12-03
4
Ilex cassine
Kakaako
Street tree
Preserved_Specimen
Allison, A.
Oahu
BISH
2021-06-01
5
Ilex cassine
Puna District, land of Olaa, along Highway 11 to Volcano 17.4 miles from Hilo
Preserved_Specimen
Herbst, D.R.
5603
Hawaii
BISH
1975-12-16
6
Ilex cassine
Pauoa Flats near junction of Pauoa Flat Trail with Aihualama Trail
Wet lowland secondary forest.
Preserved_Specimen
OED
2012061401
Oahu
BISH
2012-06-14
7
Ilex cassine
Whitmore Village
Along old dirt road about 1/2 mile from fence, on left side. Disturbed vegetation with Eucalyptus, bamboo, Ardisia, traveler's palm, Tibouchina herbacea, Clidemia hirta.
Preserved_Specimen
Keir, M.
1
Oahu
BISH
1998-12-16
8
Ilex cassine
Foster Botanic Garden
Preserved_Specimen
Lau, J.
1590
Oahu
BISH
1985-11-04
9
Ilex cassine
Lyon Arboretum, Honolulu
Preserved_Specimen
Nagata, K.M.
144
Oahu
BISH
1966-08-08
10
Ilex cassine
Foster Botanic Garden
Preserved_Specimen
Lau, J.
1590
Oahu
BISH
1985-11-04
11
Ilex cassine
Wahiawa, Public Gardens
Preserved_Specimen
Potter, C.
Oahu
BISH
1961-10-10
12
Ilex cassine
Kilauea Military Camp. Between Theater & Fire Stn
Preserved_Specimen
Stemmermann, R.L.
7016
Hawaii
BISH
1985-06-16
13
Ilex cassine
Lyon Arboretum, Honolulu
Preserved_Specimen
Nagata, K.M.
144
Oahu
BISH
1966-08-08
14
Ilex cassine
Honolulu, 3860 Old Pali Rd
Preserved_Specimen
Rock, J.F.C.
Oahu
BISH
1957-03-31
15
Ilex cassine
Wahiawa, California Ave., former H.S.P.A. nursery
Preserved_Specimen
McEldowney, G.A.
Oahu
BISH
1952-09-10
16
Ilex cassine
H.S.P.A. nursery
Preserved_Specimen
Pearsall, G.A.
Oahu
BISH
1949-06-10
17
Ilex cassine
Wahiawa, former H.S.P.A. nursery
Preserved_Specimen
Korte, K.H.
Oahu
BISH
1953-10-02
18
Ilex cassine
Wahiawa
Preserved_Specimen
Lyon, H.L.
Oahu
BISH
1955-10-12
19
Ilex cassine
Hickam Nursery
Preserved_Specimen
Pearsall, G.A.
Oahu
BISH
1949-08-12
20
Ilex cassine
Wahiawa
Preserved_Specimen
Pearsall, G.A.
Oahu
BISH
1954-09-20
21
Ilex cassine
Wahiawa, Public Gardens
Preserved_Specimen
Potter, C.
Oahu
BISH
1961-10-10
22
Ilex cassine
Wahiawa, Public Gardens
Preserved_Specimen
Potter, C.
Oahu
BISH
1961-10-10
23
Ilex cassine
Aiea, 98-851 Iliee St., residence of William and Peggy Jones
Preserved_Specimen
Lau, J.
2064
Oahu
BISH
1986-02-24
24
Ilex cassine
Naturalized along the Hilo to Volcano Hwy at about the 18 mile marker
Preserved_Specimen
Stemmermann, R.L.
6931
Hawaii
BISH
1985-04-04
25
Ilex cassine
Honolulu, Akala St., just S of King St., residential area
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voluptate velit esse cillum dolore eu fugiat nulla pariatur. Excepteur sint occaecat cupidatat
non proident, sunt in culpa qui officia deserunt mollit anim id est laborum.
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Lorem ipsum dolor sit amet, consectetur adipiscing elit, sed do eiusmod tempor incididunt ut
labore et dolore magna aliqua. Ut enim ad minim veniam, quis nostrud exercitation ullamco
laboris nisi ut aliquip ex ea commodo consequat. Duis aute irure dolor in reprehenderit in
voluptate velit esse cillum dolore eu fugiat nulla pariatur. Excepteur sint occaecat cupidatat
non proident, sunt in culpa qui officia deserunt mollit anim id est laborum.
Added By Author on 9/16/19 Photo Source
Lorem ipsum dolor sit amet, consectetur adipiscing elit, sed do eiusmod tempor incididunt ut
labore et dolore magna aliqua. Ut enim ad minim veniam, quis nostrud exercitation ullamco
laboris nisi ut aliquip ex ea commodo consequat. Duis aute irure dolor in reprehenderit in
voluptate velit esse cillum dolore eu fugiat nulla pariatur. Excepteur sint occaecat cupidatat
non proident, sunt in culpa qui officia deserunt mollit anim id est laborum.
Added By Author on 9/16/19 Photo Source
Lorem ipsum dolor sit amet, consectetur adipiscing elit, sed do eiusmod tempor incididunt ut
labore et dolore magna aliqua. Ut enim ad minim veniam, quis nostrud exercitation ullamco
laboris nisi ut aliquip ex ea commodo consequat. Duis aute irure dolor in reprehenderit in
voluptate velit esse cillum dolore eu fugiat nulla pariatur. Excepteur sint occaecat cupidatat
non proident, sunt in culpa qui officia deserunt mollit anim id est laborum.
Added By Author on 9/16/19 Photo Source
Lorem ipsum dolor sit amet, consectetur adipiscing elit, sed do eiusmod tempor incididunt ut
labore et dolore magna aliqua. Ut enim ad minim veniam, quis nostrud exercitation ullamco
laboris nisi ut aliquip ex ea commodo consequat. Duis aute irure dolor in reprehenderit in
voluptate velit esse cillum dolore eu fugiat nulla pariatur. Excepteur sint occaecat cupidatat
non proident, sunt in culpa qui officia deserunt mollit anim id est laborum.
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